Crown traits measured across the network include crown diameter and crown exposure index. To estimate crown diameter (m), the crown radius is measured from the center to the edge of the crown in eight cardinal directions, then averaged. A qualitative crown exposure index serves as a proxy for light availability is recorded following a procedure adapted from Clark & Clark (1992).
Six leaf traits are measured following the procedures of Cornelissen et al (2003): lamina size (mm2); specific leaf area (m2 kg-1); leaf thickness (μm); N concentration (mg g-1); P concentration (mg g-1); and dry matter content (mg g-1). The most recent tree census is used to randomly select 5-6 of the largest and smallest individuals of each tree species for sampling. Two to five leaves are measured for each individual. Fresh mass is recorded upon leaf removal and dry mass after drying at 60° C for 72 hrs.
Four reproductive traits are measured: dispersal mode (categorical), diaspore shape (unitless), diaspore mass (mg), and seed mass (mg). Diaspores are the unit that is dispersed by explosive force, by wind or by animals. Diaspores are dissected to isolate the embryo plus endosperms (i.e., seed). Collection of plant reproductive parts happens opportunistically and varies across sites subject to plant phenology. We attempt to collect five mature fruits from five individuals of each species, although for rare species or for those from which fruits rarely fall we collect single fruits or diaspores. Dispersal mode and shape classification follows Cornelissen et al. (2003).